This involves the player taking on the roles and detailing the stories of many other characters including Jazz, Grimlock, Starscream, the Combaticons and Cliffjumper. After some time fighting them off and Optimus personally battling Megatron, the game then jumps back some time before they left Cybertron and details the major events that occurred to get them to this point.
Intending to look for a new world to colonize, they are suddenly attacked by Megatron and his Decepticon forces. The game opens with the launching of The Ark, a massive city-ship meant to transport as many Autobots off world as possible. Because of this, it's Darker and Edgier than any other Transformers game. Now it's a race for both factions to gather what Energon and survivors they can to deal one last decisive blow to the other side before they flee the dying planet. Technically the Decepticons have won, but it's something of a Pyrrhic Victory since the war used up so much Energon and tore up most of Cybertron.
The sequel to 2010's Transformers: War for Cybertron, this game shows the tail end of the war before it left for Earth.